Tales of Beseria takes place before Zestiria as it reveals how things became to be. The story begins with three races – Malak, Humans, and Daemons. As much as possible, the three races try to coexist except during the Scarlet Hour where Daemons are at their strongest.
The story begins with the traumatic experience of Velvet when she finds herself in the middle of a Scarlet Night. While she did lose her sister 3 years prior, another Scarlet Night shatters her image of the Exorcists. This drives her to focus solely on delivering her vengeance and for exposing The Abbey. Along the way, she meets different people such as Seres, one of the first Malak. She also joins forces with Rokuro, Magilou, Laphicet – another Malak, and others. Different characters have different motives especially when they find themselves exposing how things truly are.